{"id":12130,"date":"2026-06-04T22:57:54","date_gmt":"2026-06-04T22:57:54","guid":{"rendered":"https:\/\/deliry.com\/?p=12130"},"modified":"2026-06-06T19:56:22","modified_gmt":"2026-06-06T19:56:22","slug":"rhizariens","status":"publish","type":"post","link":"https:\/\/deliry.com\/?p=12130","title":{"rendered":"Rhizariens"},"content":{"rendered":"\n<blockquote class=\"wp-block-quote is-layout-flow wp-block-quote-is-layout-flow\">\n<p><em><a href=\"https:\/\/deliry.com\/?p=9820\" data-type=\"post\" data-id=\"9820\">Naturae<\/a> <\/em>&gt; <a href=\"https:\/\/deliry.com\/?p=9825\" data-type=\"post\" data-id=\"9825\">Eucaryotes<\/a> &gt; Bicontes<\/p>\n\n\n\n<p><a href=\"https:\/\/deliry.com\/?p=9927\" data-type=\"post\" data-id=\"9927\">Protozoaires<\/a><\/p>\n<\/blockquote>\n\n\n\n<p>Les&nbsp;<strong>Rhizariens<\/strong>&nbsp;avec les&nbsp;Actinopodes&nbsp;(H\u00e9liozoaires&nbsp;et&nbsp;Radiolaires), les&nbsp;Foraminif\u00e8res&nbsp;et un groupe de&nbsp;Protophytes&nbsp;(les&nbsp;Chlorarachniophytes).<\/p>\n\n\n\n<p>Ce sont des&nbsp;Eucaryotes&nbsp;fondamentalement biflagell\u00e9s (\u00ab&nbsp;Bicontes&nbsp;\u00bb), de teinte animale (\u00abProtozoaires\u00bb ) ou v\u00e9g\u00e9tale (\u00abProtophytes\u00bb). Ils sont plac\u00e9s en parall\u00e8le de la Lign\u00e9 verte, des&nbsp;Chromoalv\u00e9ol\u00e9s&nbsp;et des&nbsp;Excavobiontes.<\/p>\n\n\n\n<p>Si des caract\u00e8res chimiques (ARNr) sont identifi\u00e9s pour associer les esp\u00e8ces de cet ensemble, il n\u2019y a pas d\u2019autre crit\u00e8re \u00e9vident pour les rassembler. L\u2019\u00e9tude de leur phylog\u00e9nie doit se poursuivre. N\u00e9anmoins ces groupes avaient d\u00e9j\u00e0 \u00e9t\u00e9 associ\u00e9s d\u00e8s la fin du XIX<sup>e<\/sup>&nbsp;si\u00e8cle dans un ensemble nomm\u00e9 \u00abRhizomon\u00e8res\u00bb (voir Railliet 1893&nbsp;: 127)\u2026 il y a donc bien des points communs particuliers (\u00e0 retrouver&nbsp;!).<\/p>\n\n\n\n<p><strong>Railliet A. 1893<\/strong>&nbsp;\u2013&nbsp;<em>Trait\u00e9 de Zoologie m\u00e9dicale et agricole.<\/em>&nbsp;Deuxi\u00e8me \u00e9dition. \u2013 Asselin &amp; Houzeau, Paris.<\/p>\n\n\n\n<p class=\"has-pale-ocean-gradient-background has-background\" style=\"font-size:25px\">Actinopodes<\/p>\n\n\n\n<p>Cet ensemble phylog\u00e9n\u00e9tique comprend les deux anciens groupes form\u00e9s par les H\u00e9liozoaires et les Radiolaires, autrefois rang\u00e9s parmi les Protozoaires, ainsi que les Acanthaires. Il est rang\u00e9 parmi les Rhizariens.<\/p>\n\n\n\n<p class=\"has-vivid-purple-color has-text-color has-link-color has-medium-font-size wp-elements-6942c1ee0a264b172e53e7da013da953\">H\u00e9liozoaires<\/p>\n\n\n\n<p>Pr\u00e9sence de pseudopodes radi\u00e9s et fins. Le test quand il est pr\u00e9sent est siliceux. Habitent essentiellement les eaux douces.<\/p>\n\n\n<div class=\"wp-block-image\">\n<figure class=\"aligncenter size-full\"><a href=\"https:\/\/deliry.com\/wp-content\/uploads\/2026\/05\/Raphidiophrys.png\"><img loading=\"lazy\" decoding=\"async\" width=\"400\" height=\"371\" src=\"https:\/\/deliry.com\/wp-content\/uploads\/2026\/05\/Raphidiophrys.png\" alt=\"\" class=\"wp-image-12730\" srcset=\"https:\/\/deliry.com\/wp-content\/uploads\/2026\/05\/Raphidiophrys.png 400w, https:\/\/deliry.com\/wp-content\/uploads\/2026\/05\/Raphidiophrys-300x278.png 300w\" sizes=\"auto, (max-width: 400px) 100vw, 400px\" \/><\/a><figcaption class=\"wp-element-caption\"><em>Raphidiophrys<\/em><br><sup>Extrait de Harmer &amp; Shipley (1906)<\/sup><\/figcaption><\/figure>\n<\/div>\n\n\n<p class=\"has-vivid-purple-color has-text-color has-link-color has-medium-font-size wp-elements-4ce79f8ba436c0363b6e0189f946b5fc\">Radiolaires<\/p>\n\n\n\n<p>Pr\u00e9sence d\u2019une chitine centrale et pseudopodes radiants et fins. Le test est siliceux. Habitent les eaux marines tant en tant que plancton que proche du fond y compris dans des zones abyssales. Les Radiolaires ont une grande importance G\u00e9ologique et sont \u00e0 l\u2019origine par bioaccumulations de roches siliceuses (g\u00e9n\u00e9ralement rouges) nomm\u00e9es&nbsp;Radiolarites.<\/p>\n\n\n<div class=\"wp-block-image\">\n<figure class=\"aligncenter size-full\"><a href=\"https:\/\/deliry.com\/wp-content\/uploads\/2026\/05\/Dorataspis.png\"><img loading=\"lazy\" decoding=\"async\" width=\"301\" height=\"297\" src=\"https:\/\/deliry.com\/wp-content\/uploads\/2026\/05\/Dorataspis.png\" alt=\"\" class=\"wp-image-12731\"\/><\/a><figcaption class=\"wp-element-caption\">Radiolaire<br><sup>d&rsquo;apr\u00e8s Haeckel<\/sup><\/figcaption><\/figure>\n<\/div>\n\n<div class=\"wp-block-image\">\n<figure class=\"aligncenter size-full is-resized\"><a href=\"https:\/\/deliry.com\/wp-content\/uploads\/2026\/05\/Radiolaires.png\"><img loading=\"lazy\" decoding=\"async\" width=\"550\" height=\"523\" src=\"https:\/\/deliry.com\/wp-content\/uploads\/2026\/05\/Radiolaires.png\" alt=\"\" class=\"wp-image-12732\" style=\"width:400px\" srcset=\"https:\/\/deliry.com\/wp-content\/uploads\/2026\/05\/Radiolaires.png 550w, https:\/\/deliry.com\/wp-content\/uploads\/2026\/05\/Radiolaires-300x285.png 300w\" sizes=\"auto, (max-width: 550px) 100vw, 550px\" \/><\/a><figcaption class=\"wp-element-caption\">Radiolaire<br><sup>Extrait de Delage &amp; H\u00e9rouard (1896)<\/sup><\/figcaption><\/figure>\n<\/div>\n\n\n<p class=\"has-vivid-purple-color has-text-color has-link-color has-medium-font-size wp-elements-7ea3dcd4d704214d65213bc3e7636bca\">Acanthaires<\/p>\n\n\n<div class=\"wp-block-image\">\n<figure class=\"alignright size-full\"><img loading=\"lazy\" decoding=\"async\" width=\"150\" height=\"149\" src=\"https:\/\/deliry.com\/wp-content\/uploads\/2026\/05\/Acanthairer.png\" alt=\"\" class=\"wp-image-12735\"\/><figcaption class=\"wp-element-caption\">Acanthaires<\/figcaption><\/figure>\n<\/div>\n\n\n<p>Les&nbsp;<strong>Acanthaires<\/strong>&nbsp;(<em>Acantharia<\/em>&nbsp;Haeckel, 1881) sont un des trois groupes des Actinopodes. Leur test est compos\u00e9 essentiellement de c\u00e9lestine qui est une association de mati\u00e8re organique avec du sulfate de strontium. Leurs axopodes sont porteurs d\u2019Algues suppos\u00e9es endosymbiotiques connues sous le nom de Zooxanthelles. Dans les faits les Acanthaires semble bloquer les divisions de leurs Algues tout en permettant la multiplication de leurs chloroplastes. Ils \u00abvolent\u00bb alors la mati\u00e8re organique produite par les Algues en quelque sorte de \u00abcytokleptie\u00bb. Dans ces termes le b\u00e9n\u00e9fice r\u00e9ciproque des deux \u00eatres vivants para\u00eet d\u00e9s\u00e9quilibr\u00e9 et en cons\u00e9quence on ne peut probablement pas parler de symbiose v\u00e9ritable. La sym\u00e9trie radi\u00e9e est particuli\u00e8rement soign\u00e9e chez les Acanthaires car les spicules sont sym\u00e9triques, stables num\u00e9riquement (dix) et ont tous la m\u00eame longueur.<\/p>\n\n\n<div class=\"wp-block-image\">\n<figure class=\"aligncenter size-full is-resized\"><a href=\"https:\/\/deliry.com\/wp-content\/uploads\/2026\/05\/Xiphacantha.png\"><img loading=\"lazy\" decoding=\"async\" width=\"350\" height=\"306\" src=\"https:\/\/deliry.com\/wp-content\/uploads\/2026\/05\/Xiphacantha.png\" alt=\"\" class=\"wp-image-12733\" style=\"aspect-ratio:1.1438273535588221;width:400px;height:auto\" srcset=\"https:\/\/deliry.com\/wp-content\/uploads\/2026\/05\/Xiphacantha.png 350w, https:\/\/deliry.com\/wp-content\/uploads\/2026\/05\/Xiphacantha-300x262.png 300w\" sizes=\"auto, (max-width: 350px) 100vw, 350px\" \/><\/a><figcaption class=\"wp-element-caption\">&nbsp;<em>Xiphacantha<\/em>&nbsp;\u2013 Squelette seulement (test)<br><sup>selon Wyville Thomson&nbsp;<em>in<\/em>&nbsp;Harmer &amp; Shipley (1906)<\/sup><\/figcaption><\/figure>\n<\/div>\n\n\n<p class=\"has-pale-ocean-gradient-background has-background\" style=\"font-size:25px\">Foraminif\u00e8res<\/p>\n\n\n<div class=\"wp-block-image\">\n<figure class=\"alignright size-full is-resized\"><a href=\"https:\/\/deliry.com\/wp-content\/uploads\/2026\/06\/Casglo.png\"><img loading=\"lazy\" decoding=\"async\" width=\"563\" height=\"599\" src=\"https:\/\/deliry.com\/wp-content\/uploads\/2026\/06\/Casglo.png\" alt=\"\" class=\"wp-image-12917\" style=\"width:300px\" srcset=\"https:\/\/deliry.com\/wp-content\/uploads\/2026\/06\/Casglo.png 563w, https:\/\/deliry.com\/wp-content\/uploads\/2026\/06\/Casglo-282x300.png 282w\" sizes=\"auto, (max-width: 563px) 100vw, 563px\" \/><\/a><figcaption class=\"wp-element-caption\"><em>Cassidulina globulosa<\/em>\u00a0\u0166 (Globig\u00e9rine) &#8211; Planche du XIX<sup>e<\/sup>\u00a0si\u00e8cle<\/figcaption><\/figure>\n<\/div>\n\n\n<p>Autrefois rang\u00e9s parmi les\u00a0Protozoaires, les\u00a0<strong>Foraminif\u00e8res<\/strong>\u00a0sont des\u00a0Eucaryotes\u00a0Bicontes\u00a0appartenant \u00e0 l\u2019ensemble des\u00a0Rhizariens.<\/p>\n\n\n\n<p>Alors que la plupart des esp\u00e8ces pr\u00e9sente un test central bien dessin\u00e9, l\u2019animal projette souvent un grand nombre de filopodes r\u00e9ticul\u00e9s (ou r\u00e9ticulopodes) maintenu par un cytosquelette tr\u00e8s dynamique. Le test est initialement de nature glycoprot\u00e9ique. La croissance se fait par addition de nouvelles loges si bien que le test de nombre de Foraminif\u00e8res ressemblent \u00e0 des coquilles microscopiques en escargot. Secondairement le test peut se calcifier o\u00f9 il peut y \u00eatre adjoint des grains de sable.<\/p>\n\n\n\n<p>Pr\u00e9sence d\u2019un test complexe form\u00e9 notamment de calcite ou form\u00e9 de grains de sables agglom\u00e9r\u00e9s et pris dans l\u2019environnement. Cet ensemble toujours actuel, est un groupe de fossiles stratigraphiques important.<\/p>\n\n\n<div class=\"wp-block-image\">\n<figure class=\"aligncenter size-full\"><img loading=\"lazy\" decoding=\"async\" width=\"331\" height=\"505\" src=\"https:\/\/deliry.com\/wp-content\/uploads\/2026\/06\/Foraminiferes.png\" alt=\"\" class=\"wp-image-12919\" srcset=\"https:\/\/deliry.com\/wp-content\/uploads\/2026\/06\/Foraminiferes.png 331w, https:\/\/deliry.com\/wp-content\/uploads\/2026\/06\/Foraminiferes-197x300.png 197w\" sizes=\"auto, (max-width: 331px) 100vw, 331px\" \/><figcaption class=\"wp-element-caption\">Tests de quelques Foraminif\u00e8res fossiles \u0166 (Harmer &amp; Shipley 1906)<br><sup>Planche reconstruite\u00a0: \u00a9\u00a9 byncsa &#8211;\u00a0Cyrille Deliry\u00a0&#8211;\u00a0<em>Histoires Naturelles<\/em><\/sup><\/figcaption><\/figure>\n<\/div>\n\n\n<hr class=\"wp-block-separator has-alpha-channel-opacity is-style-wide\"\/>\n\n\n\n<ul class=\"wp-block-list\">\n<li><strong>Delage Y. &amp; H\u00e9rouard E. 1896<\/strong>&nbsp;&#8211;&nbsp;<em>Trait\u00e9 de Zoologie concr\u00e8te.Tome I. La cellule et les Protozoaires.<\/em>&nbsp;&#8211; Reinwald,, Schleicher, Paris.<\/li>\n\n\n\n<li><strong>Harmer S.F. &amp; Shipley A.E. 1895-1909<\/strong>&nbsp;&#8211;&nbsp;<em>The Cambridge natural history.<\/em>&nbsp;&#8211; London.<\/li>\n\n\n\n<li><strong>Lecointre G. &amp; Le Guyader H. 2001<\/strong>&nbsp;&#8211;&nbsp;<em>Classification phylog\u00e9n\u00e9tique du vivant.<\/em>&nbsp;&#8211; Belin, Paris.<\/li>\n<\/ul>\n","protected":false},"excerpt":{"rendered":"<p>Naturae &gt; Eucaryotes &gt; Bicontes Protozoaires Les&nbsp;Rhizariens&nbsp;avec les&nbsp;Actinopodes&nbsp;(H\u00e9liozoaires&nbsp;et&nbsp;Radiolaires), les&nbsp;Foraminif\u00e8res&nbsp;et un &hellip; <\/p>\n","protected":false},"author":1,"featured_media":12735,"comment_status":"closed","ping_status":"closed","sticky":false,"template":"","format":"standard","meta":{"footnotes":""},"categories":[535,1,543,563],"tags":[],"class_list":["post-12130","post","type-post","status-publish","format-standard","has-post-thumbnail","hentry","category-eucaryotes","category-maintenance","category-protozoaires","category-rhizariens"],"_links":{"self":[{"href":"https:\/\/deliry.com\/index.php?rest_route=\/wp\/v2\/posts\/12130","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/deliry.com\/index.php?rest_route=\/wp\/v2\/posts"}],"about":[{"href":"https:\/\/deliry.com\/index.php?rest_route=\/wp\/v2\/types\/post"}],"author":[{"embeddable":true,"href":"https:\/\/deliry.com\/index.php?rest_route=\/wp\/v2\/users\/1"}],"replies":[{"embeddable":true,"href":"https:\/\/deliry.com\/index.php?rest_route=%2Fwp%2Fv2%2Fcomments&post=12130"}],"version-history":[{"count":5,"href":"https:\/\/deliry.com\/index.php?rest_route=\/wp\/v2\/posts\/12130\/revisions"}],"predecessor-version":[{"id":12920,"href":"https:\/\/deliry.com\/index.php?rest_route=\/wp\/v2\/posts\/12130\/revisions\/12920"}],"wp:featuredmedia":[{"embeddable":true,"href":"https:\/\/deliry.com\/index.php?rest_route=\/wp\/v2\/media\/12735"}],"wp:attachment":[{"href":"https:\/\/deliry.com\/index.php?rest_route=%2Fwp%2Fv2%2Fmedia&parent=12130"}],"wp:term":[{"taxonomy":"category","embeddable":true,"href":"https:\/\/deliry.com\/index.php?rest_route=%2Fwp%2Fv2%2Fcategories&post=12130"},{"taxonomy":"post_tag","embeddable":true,"href":"https:\/\/deliry.com\/index.php?rest_route=%2Fwp%2Fv2%2Ftags&post=12130"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}